A second update on mapping the human genetic architecture of COVID-19


Kanai M. Andrews S.J. Cordioli M. Stevens C. Neale B.M. Daly M. Ganna A. Pathak G.A. Iwasaki A. Karjalainen J. Mehtonen J. Pirinen M. Chwialkowska K. Trankiem A. Balaconis M.K. Veerapen K. Wolford B.N. Ahmad H.F. von Hohenstaufen Puoti K.A. Boer C. Boua P.R. Butler-Laporte G. Cadilla C.L. Colombo F. Douillard V. Dueker N. Dutta A.K. El-Sherbiny Y.M. Eltoukhy M.M. Esmaeeli S. Faucon A. Fave M.-J. Cadenas I.F. Francescatto M. Francioli L. Franke L. Fuentes M. Durán R.G. Cabrero D.G. Harry E.N. Jansen P. Szentpéteri J.L. Kaja E. Kirk C. Kousathanas A. Krieger J.E. Patel S.K. Lemaçon A. Limou S. Lió P. Marouli E. Marttila M.M. Medina-Gómez C. Michaeli Y. Migeotte I. Mondal S. Moreno-Estrada A. Moya L. Nakanishi T. Nasir J. Pasko D. Pearson N.M. Pereira A.C. Priest J. Prijatelj V. Prokic I. Teumer A. Várnai R. Romero-Gómez M. Roos C. Rosenfeld J. Ruolin L. Schulte E.C. Schurmann C. Sedaghati-khayat B. Shaheen D. Shivanathan I. Sipeky C. Sirui Z. Striano P. Tanigawa Y. Remesal A.U. 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Krishnamurthy V. Diaba C. John L. Lim L. Jha R. Egan J. Felton T. Glasgow S. Padden G. Choudhr O. Bradley-Potts J. Moss S. Lingeswaran S. Alexander P. Brandwood C. Fiouni S. Ward L. Allen S. Shaw J. Smith C. Adanini O. Collins R. Msiska M. Ofori L. Bhatia N. Dolan H. Brunton M. Caterson J. Coles H. Keating L. Tilney E. Jacques N. Frise M. Armistead J. Bartley S. Bhuie P. Rai S. Tomkova G. Greer S. Shuker K. Tridente A. Dobson E. Hunt J. Tully R. Dearden J. Drummond A. Kamath P. Bullock E. Mulcahy M. Munt S. O’Connor G. Philbin J. Rishton C. Scott C. Winnard S. Hasni N. Gascoyne R. Hawes J. Pritchard K. Stevenson L. Whileman A. Beavis S. Bishop L. Cart C. Dale K. Kelly-Baxter M. Mendelski A. Moakes E. Smith R. Woodward J. Wright S. Allan A. Botello A. Liew J. Medhora J. Trumper E. Savage F. Scott T. Place M. Kaye C. Benyon S. Marriott S. Park L. Quinn H. Skyes D. Zitter L. Baines K. Gordon E. Keenan S. Pitt A. Duffy K. Ireland J. Semple G. Turner L. Cathcart S. Rimmer D. Puxty A. Puxty K. Hurst A. Miller J. Speirs S. Walker L. Bradshaw Z. Brown J. Melling S. Preston S. Slawson N. Warden S. Beasley A. Stoddard E. Benham L. Cupitt J. Caswell M. Elawamy L. Wignall A. Roberts B. Golding H. Leggett S. Male M. Marani M. Prager K. Williams T. Golder K. Jones O. Cusack R. Bolger C. Burnish R. Carter M. Jackson S. Salmon K. Biss J. Aquino M. Croft M. Frost V. White I. Govender K. Webb N. Stapleton L. Wells C. Nikitas N. Sanchez-Rodriguez A. Spencer K. Stowe B. Izzard Y. Poole M. Monnery S. Trotman S. Beech V. Combes E. Joefield T. Covernton P. Savage S. Woodward E. Camsooksai J. Reschreiter H. Barclay C. DeAth Y. Dube J. Humphrey C. Jenkins S. Langridge E. Milne R. Wadams B. Woolcock M. Brett M. Digby B. Gemmell L. Hornsby J. MacGoey P. O’Neil P. Price R. Sundaram R. Rodden N. Thomson N. Rooney K.
7 September 2023Nature Research

Nature
2023#621Issue 7977E7 - E26 pp.





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Broad Institute of MIT and Harvard, Cambridge, MA, United States
University of California San Francisco, San Francisco, CA, United States
Institute for Molecular Medicine Finland (FIMM), University of Helsinki, Helsinki, Finland
Analytic and Translational Genetics Unit, Massachusetts General Hospital, Boston, MA, United States
Department of Psychiatry, School of Medicine, Yale University, New Haven, CT, United States
Veteran Affairs Connecticut Healthcare System, West Haven, CT, United States
Yale University, New Haven, CT, United States
Centre for Bioinformatics and Data Analysis, Medical University of Bialystok, Bialystok, Poland
Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, MI, United States
Faculty of Industrial Sciences and Technology, Universiti Malaysia Pahang, Pahang, Malaysia
Icahn School of Medicine at Mount Sinai, New York, NY, United States
Genolier Innovation Network and Hub, Swiss Medical Network, Genolier Healthcare Campus, Genolier, Switzerland
Immediate, Independent Multidisciplinary Med & Biotech Research Association, Milan, Italy
Clinical Research Unit of Nanoro, Institut de Recherche en Sciences de la Santé, Nanoro, Burkina Faso
Sydney Brenner Institute for Molecular Biosciences, University of the Witwatersrand, Johannesburg, South Africa
McGill University, Montreal, QC, Canada
University of Peurto Rico, San Juan, Puerto Rico
Institute for Biomedical Technologies, National Research Council, Milan, Italy
University of Miami, Miami, FL, United States
Department of Biochemistry, All India Institute of Medical Sciences, Kalyani, India
Department of Biosciences, School of Science and Technology, Nottingham Trent University, Nottingham, United Kingdom
Department of Clinical Pathology, Faculty of Medicine, Mansoura University, Mansoura, Egypt
Department of Clinical Pathology, Faculty of Medicine, Tanta University, Tanta, Egypt
Neuron23, San Francisco, CA, United States
Vanderbilt University, Nasvhille, TN, United States
Stroke Pharmacogenomics and Genetics, Institut d’Investigació Biomèdica Sant Pau (IIBSANT PAU), Barcelona, Spain
Department of Medicine, Surgery and Health Sciences, University of Trieste, Trieste, Italy
Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, United States
Department of Genetics, University Medical Centre Groningen, University of Groningen, Groningen, Netherlands
Departamento de Tecnología Médica, Facultad de Ciencias de la Salud, Universidad de Tarapacá, Arica, Chile
King’s College London, London, United Kingdom
Naina Tech, New York, NY, United States
Vrije Universiteit Amsterdam, Amsterdam, Netherlands
Medical School, University of Pécs, Pécs, Hungary
MNM Diagnostics, Poznan, Poland
Genomics England, London, United Kingdom
Heart Institute (InCor), University of Sao Paulo Medical School, Sao Paulo, Brazil
Department of Physiology, All India Institute of Medical Sciences, Kalyani, India
Montreal Heart Institute, Montreal, QC, Canada
Nantes Université, Ecole Centrale Nantes, INSERM, Center for Research in Transplantation and Translational Immunology, Nantes, France
Department of Computer Science and Technology, University of Cambridge, Cambridge, United Kingdom
William Harvey Research Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London, United Kingdom
University of Helsinki, Helsinki, Finland
Department of Internal Medicine, Erasmus MC, University Medical Center Rotterdam, Rotterdam, Netherlands
Fonds de la Recherche Scientifique (FNRS) & Centre de Génétique Humaine, Hôpital Erasme, Université Libre de Bruxelles, Brussels, Belgium
Department of Genetics, Hôpital Erasme, Université Libre de Bruxelles, Brussels, Belgium
All India Institute of Medical Sciences Kalyani, Kalyani, India
National Laboratory of Genomics for Biodiversity, CINVESTAV, Guanajuato, Mexico
Queensland University of Technology, Brisbane, QLD, Australia
Department of Human Genetics, McGill University, Montreal, QC, Canada
Lady Davis Institute, Jewish General Hospital, McGill University, Montreal, QC, Canada
Kyoto-McGill International Collaborative School in Genomic Medicine, Graduate School of Medicine, Kyoto University, Kyoto, Japan
Japan Society for the Promotion of Science, Tokyo, Japan
Division of Life Sciences, University of Northampton, Northampton, United Kingdom
Root Deep, Boston, MA, United States
Instituto do Coração, Sao Paulo, Brazil
Stanford University, Stanford, CA, United States
Department of Epidemiology, Erasmus MC, University Medical Center Rotterdam, Rotterdam, Netherlands
Institute for Community Medicine, University Medicine Greifswald, Greifswald, Germany
GI and Liver Section, Virgen del Rocío University Hospital and Department of Medicine, University of Seville, Institute of Biomedicine of Seville, Seville, Spain
CIBEREHD, Instituto de Salud Carlos III (ISCIII), Seville, Spain
Department of Health, Traunstein, Germany
Rutgers Cancer Institute of New Jersey, New Brunswick, NJ, United States
Erasmus Medical Center, Rotterdam, Netherlands
Institute of Psychiatric Phenomics & Genomics, University of Munich, Munich, Germany
Department of Psychiatry, University of Munich, Munich, Germany
Institute of Virology, Technical University of Munich/Helmholtz Munich, Munich, Germany
Institute of Human Genetics, University Hospital, Faculty of Medicine, University of Bonn, Bonn, Germany
Department of Psychiatry, University Hospital, Faculty of Medicine, University of Bonn, Bonn, Germany
Digital Health Center, Hasso Plattner Institute, University of Potsdam, Potsdam, Germany
Institute of Biomedicine and Cancer Research Laboratories, Western Cancer Centre FICAN West, University of Turku, Turku, Finland
IRCCS Istituto Giannina Gaslini, Genova, Italy
Department of Neurosciences, Rehabilitation, Ophthalmology, Genetics, Maternal and Child Health, University of Genova, Genova, Italy
Department of Biomedical Data Science, Stanford University, Stanford, CA, United States
Fundación Pública Andaluza para la Gestión de la Investigación en Salud de Sevilla, Seville, Spain
Department of Pediatrics, Stanford University, Stanford, CA, United States
University Medical Center Utrecht, Utrecht, Netherlands
Instituto de Investigación Interdisciplinaria & Escuela de Medicina, Universidad de Talca, Talca, Chile
Programa de Genética Humana, ICBM and Chile Departamento de Oncología Básico Clínica, Universidad de Chile, Talca, Chile
Department of Statistical Genetics, Osaka University Graduate School of Medicine, Osaka, Japan
Department of Internal Medicine, International Islamic University Malaysia, Pahang, Malaysia
Centro para el Desarrollo de la Investigación Científica (CEDIC), Asunción, Paraguay
23andMe, Sunnyvale, CA, United States
Washington University School Of Medicine, St Louis, MO, United States
Department of Psychological & Brain Sciences, Washingon University in St Louis, St Louis, MO, United States
Institute of Biological Psychiatry, Mental Health Centre Sct Hans, Mental Health Services Capital Region of Denmark, Copenhagen, Denmark
iPSYCH, The Lundbeck Foundation Initiative for Integrative Psychiatric Research, Aarhus, Denmark
Laureate Institute for Brain Research, Tulsa, OK, United States
Department of Radiology, UCSD, San Diego, CA, United States
Nork Infection Clinical Hospital, Yerevan, Armenia
Institute of Molecular Biology NAS RA, Yerevan, Armenia
National Research Center for Cardiac Surgery, Astana, Kazakhstan
City Infectious Disease Center, Astana, Kazakhstan
National Center for Biotechnology, Astana, Kazakhstan
Health Department of Astana, Astana, Kazakhstan
MRC Integrative Epidemiology Unit, University of Bristol, Bristol, United Kingdom
Division of Psychitary, University of Edinburgh, Edinburgh, United Kingdom
Institute for Molecular Medicine Finland (FIMM), Helsinki, Finland
University of Liege, GIGA-Institute, Liege, Belgium
CHC Mont-Légia, Liege, Belgium
BHUL (Liège Biobank), CHU of Liège, Liege, Belgium
Institute for Molecular Medicine Finland, University of Helsinki, Helsinki, Finland
Analytic & Translational Genetics Unit, Massachusetts General Hospital, Boston, MA, United States
Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, United States
CHU of Liege, Liege, Belgium
University of Liege, Liege, Belgium
Centre de Génétique Humaine, Hôpital Erasme, Université Libre de Bruxelles, Brussels, Belgium
Service de Médecine Interne, Hôpital Erasme, Université Libre de Bruxelles, Brussels, Belgium
McGill Genome Centre and Department of Human Genetics, McGill University, Montréal, QC, Canada
Department of Human Genetics, Epidemiology, Biostatistics and Occupational Health, McGill University, Montreal, QC, Canada
Department of Twin Research, King’s College London, London, United Kingdom
The Meakins-Christie Laboratories at the Research Institute of the McGill University Heath Centre Research Institute and Department of Medicine, Faculty of Medicine, McGill University, Montreal, QC, Canada
Research Centre of the Centre Hospitalier de l’Université de Montréal (CRCHUM), Montreal, QC, Canada
Centre Hospitalier de l’Université de Montréal (CHUM), Montreal, QC, Canada
Department of Epidemiology, Biostatistics and Occupational Health, McGill University, Montreal, QC, Canada
Department of Emergency Medicine, McGill University, Montreal, QC, Canada
Emergency Department, Jewish General Hospital, McGill University, Montreal, QC, Canada
McGill AIDS Centre, Department of Microbiology and Immunology, Lady Davis Institute for Medical Research, Jewish General Hospital, McGill University, Montreal, QC, Canada
Division of Infectious Diseases, Lausanne University Hospital and University of Lausanne, Lausanne, Switzerland
Research Centre of the Centre Hospitalier de l’Université de Montréal and Université de Montréal, Montreal, QC, Canada
School of Rehabilitation, Faculty of Medicine and Health Sciences, Université de Sherbrooke, Sherbrooke, Canada
Department of Medicine, Faculty of Medicine, Université de Montréal, Montreal, QC, Canada
Center for Immunity, Inflammation and Infectious Diseases, Montreal Clinical Research Institute (IRCM), Montreal, QC, Canada
Génome Québec, Montreal, QC, Canada
Département de Microbiologie et Infectiologie, Université de Sherbrooke, Sherbrooke, QC, Canada
Département de Médecine, Service d’Infectiologie, Centre de Recherche Clinique du Centre Hospitalier Universitaire de Sherbrooke, Sherbrooke, QC, Canada
Center for Clinical Epidemiology, Lady Davis Institute, Jewish General Hospital, Montreal, QC, Canada
Department of Neurology and Neurosurgery, McGill University, Montreal, QC, Canada
Centre Intégré Universitaire de Santé et de Services Sociaux (CIUSSS) du Saguenay–Lac-Saint-Jean, Saguenay, QC, Canada
Department of Pharmacology-Physiology, Medicine and Health Sciences Faculty, Université de Sherbrooke, Sherbrooke, QC, Canada
Division of Respiratory Medicine, Department of Pediatrics, Sainte-Justine University Hospital Center, University of Montreal, Montreal, QC, Canada
Centre of Genomics and Policy, McGill University, Montreal, QC, Canada
Division of Genetic Medicine, Department of Medicine, Vanderbilt University Medical Center, Nashville, TN, United States
Vanderbilt Genetics Institute, Vanderbilt University Medical Center, Nashville, TN, United States
Institute of Human Genetics, University Hospital Bonn, Medical Faculty University of Bonn, Bonn, Germany
Institute of Genomic Statistics and Bioinformatics, University Hospital Bonn, Medical Faculty University of Bonn, Bonn, Germany
Department of Gastroenterology, Hepatology and Infectious Diseases, University Hospital Duesseldorf, Medical Faculty Heinrich Heine University, Duesseldorf, Germany
Department of Gastroenterology, Hepatology and Infectiology, University Hospital Magdeburg, Magdeburg, Germany
Clinic for Cardiology, Angiology and Internal Intensive Medicine, Medical Clinic I, RWTH Aachen University, Aachen, Germany
Department of Pneumology and Intensive Care Medicine, Faculty of Medicine, RWTH Aachen University, Aachen, Germany
Department of Pneumology, Hannover Medical School, Hannover, Germany
Department of Gastroenterology, Hepatology and Endocrinology, Hannover Medical School, Hannover, Germany
Hannover Unified Biobank, Hannover Medical School, Hannover, Germany
Department I of Internal Medicine, Faculty of Medicine and University Hospital of Cologne, University of Cologne, Cologne, Germany
Center for Molecular Medicine Cologne (CMMC), University of Cologne, Cologne, Germany
German Center for Infection Research (DZIF), partner site Bonn-Cologne, Cologne, Germany
Institute for Human Genetics and Genomic Medicine, Medical Faculty, RWTH Aachen University, Aachen, Germany
Department of Anesthesiology and Intensive Care Medicine, University Hospital Essen, University Duisburg-Essen, Essen, Germany
Department of Child and Adolescent Psychiatry, University Hospital Essen, University of Duisburg-Essen, Essen, Germany
Department of Infectious Diseases, University Hospital Essen, University Duisburg-Essen, Essen, Germany
Department of Pneumology, Allergology and Respiratory Medicine, University Hospital Saarland, Homburg, Germany
Center of Human and Molecular Biology, Department of Human Genetics, University Hospital Saarland, Homburg, Germany
Department of Genetics & Epigenetics, Saarland University, Saarbrücken, Germany
Institute of Innate Immunity, University Hospital Bonn, Bonn, Germany
University of Toronto, Toronto, ON, Canada
Dalla Lana School of Public Health, University of Toronto, Toronto, ON, Canada
Lunenfeld-Tanenbaum Research Institute, Sinai Health, Toronto, ON, Canada
Simon Fraser University, Burnaby, BC, Canada
University of Waterloo, Waterloo, ON, Canada
The Hospital for Sick Children, Toronto, ON, Canada
Queen’s University, Kingston, ON, Canada
Mount Sinai Hospital, Sinai Health, Toronto, ON, Canada
St Michael’s Hospital, Unity Health Toronto, Toronto, ON, Canada
Princess Margaret Cancer Center, University Health Network, Toronto, ON, Canada
Ontario Institute for Cancer Research, Toronto, ON, Canada
Dynacare Laboratory, Toronto, ON, Canada
William Osler Health System, Brampton, ON, Canada
University Health Network, Toronto, ON, Canada
Mackenzie Health, Toronto, ON, Canada
Women’s College Hospital, Toronto, ON, Canada
University of British Columbia, Vancouver, BC, Canada
BC Children’s Hospital, Vancouver, BC, Canada
St Paul’s Hospital, Vancouver, BC, Canada
University of Calgary, Calgary, AB, Canada
Alberta Children’s Hospital, Calgary, AB, Canada
British Columbia Cancer Research Centre, Vancouver, BC, Canada
Toronto General Hospital, Toronto, ON, Canada
Division of Infectious Diseases, Department of Medicine, University of Ottawa, Ottawa, ON, Canada
Department of Biochemistry, Microbiology and Immunology, University of Ottawa Centre of Infection, Immunity and Inflammation (CI3), Ottawa, ON, Canada
The Ottawa Hospital Research Institute, Ottawa, ON, Canada
University of Montreal, Montreat, QC, Canada
Ste-Justine University Health Centre, Montreal, QC, Canada
University of Toronto Ontario, Toronto, ON, Canada
MitogenDx/Eve Technologies, Toronto, ON, Canada
Genome Sciences Centre, BC Cancer, Vancouver, BC, Canada
Montreal Children’s Hospital, Montreal, QC, Canada
London Health Sciences Centre, London, ON, Canada
Kidney and Metabolism Program, St Michael’s Hospital, Unity Health Toronto, Toronto, ON, Canada
Sinai Health, Mount Sinai Hospital, Toronto, ON, Canada
Baycrest Health Sciences, Toronto, ON, Canada
University of Colorado, Anschutz Medical Campus, Aurora, CO, United States
Institute of Psychiatric Phenomics and Genomics (IPPG), University Hospital, LMU Munich, Munich, Germany
Department of Genetic Epidemiology in Psychiatry, Central Institute of Mental Health, Medical Faculty Mannheim, University of Heidelberg, Mannheim, Germany
Department of Psychiatry and Behavioral Sciences, Johns Hopkins University, Baltimore, MD, United States
Department of Psychiatry and Behavioral Sciences, SUNY Upstate Medical University, Syracuse, NY, United States
Department of Psychiatry and Psychotherapy, University Hospital, LMU Munich, Munich, Germany
Institute of Virology, Technical University Munich/Helmholtz Zentrum München, Munich, Germany
Department of Psychiatry and Psychotherapy, University Hospital Bonn, Medical Faculty University of Bonn, Bonn, Germany
CIBER de Salud Mental, Instituto de Salud Carlos III, Barcelona, Spain
Munich Clinic Schwabing, Academic Teaching Hospital, Ludwig-Maximilians University of Munich (LMU), Munich, Germany
Department of Internal Medicine II, University Hospital rechts der Isar, Technical University of Munich, School of Medicine, Munich, Germany
Institute of Clinical Chemistry and Pathobiochemistry, School of Medicine, Technical University of Munich, Munich, Germany
TranslaTUM, Center for Translational Cancer Research, Technical University of Munich, Munich, Germany
Department of Psychiatry and Psychotherapy, University Medical Center Goettingen, Goettingen, Germany
German Center for Neurodegenerative Diseases (DZNE), Goettingen, Germany
Neurosciences and Signaling Group, Institute of Biomedicine (iBiMED), Department of Medical Sciences, University of Aveiro, Agra do Crasto, Aveiro, Portugal
Munich Site of the German Center for Infection Research (DZIF), Munich, Germany
Institute of Computational Biology, Helmholtz Center Munich, Oberschleissheim, Germany
Department of Hygiene and Epidemiology, University of Ioannina School of Medicine, Ioannina, Greece
1st Department of Internal Medicine and Infectious Diseases Unit, University Hospital of Ioannina, Ioannina, Greece
Department of Epidemiology and Biostatistics, School of Public Health, Imperial College London, London, United Kingdom
Center for Evidence-Based Medicine, Department of Health Services, Policy and Practice, School of Public Health, Brown University, Providence, RI, United States
University of Ioannina School of Medicine, Ioannina, Greece
Centre for Systems Biology, Biomedical Research Foundation of the Academy of Athens, Athens, Greece
Institute of Clinical Molecular Biology, Christian-Albrechts-University, Kiel, Germany
Novo Nordisk Foundation Center for Protein Research, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark
Institut de Biotecnologia i de Biomedicina, Universitat Autònoma de Barcelona, Barcelona, Spain
ICREA, Barcelona, Spain
Institute of Biotechnology, Life Science Centre, Vilnius University, Vilnius, Lithuania
Research Group for Evolutionary Immunogenomics, Max Planck Institute for Evolutionary Biology, Plön, Germany
Research Unit for Evolutionary Immunogenomics, Department of Biology, University of Hamburg, Hamburg, Germany
Department of Gastroenterology, Hospital Universitario Ramón y Cajal, University of Alcalá, Instituto Ramón y Cajal de Investigación Sanitaria (IRYCIS), Madrid, Spain
Centro de Investigación Biomédica en Red en Enfermedades Hepáticas y Digestivas (CIBEREHD), Instituto de Salud Carlos III (ISCIII), Madrid, Spain
Vall d’Hebron Institut de Recerca (VHIR), Vall d’Hebron Hospital Universitari, Barcelona, Spain
Fondazione IRCCS Ca’ Granda Ospedale Maggiore Policlinico, Milan, Italy
Department of Medicine, Møre & Romsdal Hospital Trust, Ålesund, Norway Geminicenter for Sepsis Research, Institute of Circulation and Medical Imaging (ISB), NTNU, Trondheim, Norway
Microbiology Unit, Hospital Universitario Clinico San Cecilio, Granada, Spain
Ibs.Granada Instituto de Investigación Biosanitaria, Granada, Spain
Klinik für Innere Medizin I, Universitätsklinikum Schleswig-Holstein, Campus Kiel, Kiel, Germany
Emergency Department, University Hospital Regensburg, Regensburg, Germany
Department for Infectious Diseases and Infection Control, University Hospital Regensburg, Regensburg, Germany
Department of Medicine I, Gastroenterology, Hepatology and Endocrinology, Medical University of Innsbruck, Innsbruck, Austria
Christian Doppler Laboratory of Iron and Phosphate Biology at the Department of Medicine I, Medical University of Innsbruck, Innsbruck, Austria
Institute of Clinical Medicine, University of Oslo, Oslo, Norway
Department of Microbiology, Oslo University Hospital, Oslo, Norway
Hospital Clinic, University of Barcelona and IDIBAPS, Barcelona, Spain
European Foundation for the Study of Chronic Liver Failure (EF-CLIF), Barcelona, Spain
GCAT—Genomes for Life, Germans Trias i Pujol Health Sciences Research Institute (IGTP), Badalona, Spain
Genomes for Life—GCAT Laboratory, Germans Trias i Pujol Research Institute (IGTP), Spainermans Trias i Pujol Research Institute (IGTP), Badalona, Spain
University of Milan, Milan, Italy
Department of Liver and Gastrointestinal Diseases, Biodonostia Health Research Institute, Donostia University Hospital, University of the Basque Country (UPV/EHU), CIBEREHD, Ikerbasque, San Sebastian, Spain
Digestive Diseases Unit, Virgen del Rocio University Hospital, Institute of Biomedicine of Seville, University of Seville, Seville, Spain
University of Sevilla, Seville, Spain
Instituto de Biomedicina de Sevilla (IBIS), Seville, Spain
Hospital Universitario Virgen del Rocío de Seville, Seville, Spain
Universitat Autònoma de Barcelona, Bellatera, Spain
Liver Unit, Department of Internal Medicine, Hospital Universitari Vall d’Hebron, Vall d’Hebron Barcelona Hospital Campus, Barcelona, Spain
Gastrointestinal Genetics Lab, CIC bioGUNE—BRTA, Derio, Spain
Ikerbasque, Basque Foundation for Science, Bilbao, Spain
Department of Liver and Gastrointestinal Diseases, Biodonostia Health Research Institute—Donostia University Hospital, University of the Basque Country (UPV/EHU), CIBEREHD, Ikerbasque, San Sebastian, Spain Ikerbasque, Basque Foundation for Science, Bilbao, Spain
Department I of Internal Medicine, Faculty of Medicine and University Hospital Cologne, University of Cologne, Cologne, Germany
University of Cologne, Faculty of Medicine and University Hospital Cologne, Cologne Excellence Cluster on Cellular Stress Responses in Aging-Associated Diseases (CECAD), Cologne, Germany
European Reference Network on Hepatological Diseases (ERN RARE-LIVER), Fondazione IRCCS San Gerardo dei Tintori, Monza, Italy
Division of Gastroenterology, Center for Autoimmune Liver Diseases, School of Medicine and Surgery, University of Milano-Bicocca, Milan, Italy
Genomes for Life—GCAT lab, Germans Trias i Pujol Research Institute (IGTP), Badalona, Spain
IRCCS Humanitas Research Hospital, Milan, Italy
Department of Biomedical Sciences, Humanitas University, Pieve Emanuele, Milan, Italy
Zentrum für Humangenetik Regensburg, Regensburg, Germany
University Hospital Schleswig-Holstein (UKSH), Campus Kiel, Kiel, Germany
Section for Gastroenterology, Department of Transplantation Medicine, Division for Cancer Medicine, Surgery and Transplantation, Oslo University Hospital Rikshospitalet, Oslo, Norway
Research Institute for Internal Medicine, Division of Surgery, Inflammatory Diseases and Transplantation, Oslo University Hospital Rikshospitalet and University of Oslo, Oslo, Norway
Norwegian PSC Research Center, Department of Transplantation Medicine, Division of Surgery, Inflammatory Diseases and Transplantation, Oslo University Hospital Rikshospitalet, Oslo, Norway
Private University in the Principality of Liechtenstein, Triesen, Liechtenstein
Division of Rheumatology, Inflammation and Immunity, Brigham and Women’s Hospital and Harvard Medical School, Boston, MA, United States
Division of Genetics, Department of Medicine, Brigham and Women’s Hospital, Boston, MA, United States
Department of Biomedical Informatics, Harvard Medical School, Boston, MA, United States
Center for Data Sciences, Brigham and Women’s Hospital, Boston, MA, United States
Institut de Biotecnologia i de Biomedicina, Universitat Autònoma de Barcelona, Bellaterra, Barcelona, Spain
Institute for Medical Microbiology and Hospital Epidemiology, Hannover Medical School, Hannover, Germany
Randaberg Municipality, Randaberg, Norway
Department of Quality and Health Technology, Faculty of Health Sciences, University of Stavanger, Stavanger, Norway
University Hospital Schleswig-Holstein, Campus Kiel, Kiel, Germany
Research Department, Stavanger University Hospital, Stavanger, Norway
Department of Genetics and Bioinformatics (HDGB), Division of Health Data and Digitalization, Norwegian Institute of Public Health, Oslo, Norway
Centre for Genetics and Genomics Versus Arthritis, Centre for Musculoskeletal Research, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom
Department of Intensive Care, Hospital Universitario Ramón y Cajal, Instituto Ramón y Cajal de Investigación Sanitaria (IRYCIS), University of Alcalá, Madrid, Spain
Clinical Biochemistry Department, Donostialdea Integrated Health Organisation, Osakidetza Basque Health Service, San Sebastian, Spain
Research Center and Memory Clinic, Ace Alzheimer Center Barcelona—Universitat Internacional de Catalunya, Barcelona, Spain
CIBERNED, Network Center for Biomedical Research in Neurodegenerative Diseases, National Institute of Health Carlos III, Madrid, Spain
Department of Biochemistry, University Hospital Vall d’Hebron, Barcelona, Spain
Department of Acute Medicine, Oslo University Hospital, Oslo, Norway
Division of Neurogenetics and Molecular Psychiatry, Department of Psychiatry and Psychotherapy, Faculty of Medicine and University Hospital Cologne, University of Cologne, Cologne, Germany
Department of Neurodegenerative Diseases and Geriatric Psychiatry, Medical Faculty, University Hospital Bonn, Bonn, Germany
German Center for Neurodegenerative Diseases (DZNE), Bonn, Germany
Department of Psychiatry, Glenn Biggs Institute for Alzheimer’s and Neurodegenerative Diseases, San Antonio, TX, United States
Department of Anesthesiology and Intensive Care, University Hospital of North Norway, Tromsø, Norway
Pediatrics and Tettamanti Center, Fondazione IRCCS San Gerardo dei Tintori and School of Medicine and Surgery, University of Milano-Bicocca, Monza, Italy
Biochemistry Department, Echevarne Laboratory, Sant Cugat del Vallés, Barcelona, Spain
Centre for Multidisciplinary Research in Health Science (MACH), University of Milan, Milan, Italy
Gastroenterology Unit, Fondazione IRCCS Casa Sollievo della Sofferenza, San Giovanni Rotondo, Italy
Microbiology Unit, Fondazione IRCCS San Gerardo, Monza, Italy
Department of Infectious Diseases, Oslo University Hospital, Oslo, Norway
Humanitas Gavazzeni-Castelli, Bergamo, Italy
Microbiology Department, Hospital Universitari Vall d’Hebron, Barcelona, Spain
Department of Respiratory Diseases, Hospital Universitario Ramón y Cajal, Instituto Ramón y Cajal de Investigación Sanitaria (IRYCIS), University of Alcalá, Centro de Investigación Biomédica en Red en Enfermedades Respiratorias (CIBERES), Madrid, Spain
Department of Respiratory Medicine and Allergology, University Hospital, Goethe University, Frankfurt am Main, Germany
Centro de Investigación Biomédica en Red de Enfermedades Respiratorias (CIBERES), Madrid, Spain
Centro de Investigación Biomédica en Red de Epidemiología y Salud Pública (CIBERESP), Madrid, Spain
Consejo Superior de Investigaciones Científicas, Seville, Spain
Department of Infectious Diseases, Hospital Universitario Ramón y Cajal, Instituto Ramón y Cajal de Investigación Sanitaria (IRYCIS), University of Alcalá, Madrid, Spain
Department of Transfusion Medicine and Haematology Laboratory, San Gerardo Hospital, Monza, Italy
Respiratory Medicine & International Health, University of Lübeck, Lübeck, Germany
Division of Clinical Infectious Diseases, Research Center Borstel, Borstel, Germany
Clinical Tuberculosis Unit, German Center for Infection Research (DZIF), Borstel, Germany
Internal Medicine Department, Fondazione IRCCS Ca’ Granda Ospedale Maggiore Policlinico, Milan, Italy
Humanitas Clinical and Research Center, IRCCS, Milan, Italy
Respiratory Service, Basurto University Hospital, Osakidetza Basque Health Service, Bilbao, Spain
Department of Clinical and Molecular Medicine, Faculty of Medicine and Health Sciences, Norwegian University of Science and Technology, Trondheim, Norway
Department of Medicine, Møre & Romsdal Hospital Trust, Ålesund, Norway
Department of Internal Medicine II, Medical University of Innsbruck, Innsbruck, Austria
Department of Anesthesiology and Critical Care, Hospital Universitario Ramón y Cajal, Instituto Ramón y Cajal de Investigación Sanitaria (IRYCIS), University of Alcalá, Madrid, Spain
Immunohematology Department, Banc de Sang i Teixits, Autonomous University of Barcelona, Barcelona, Spain
European Reference Network on Hepatological Diseases (ERN RARE-LIVER), San Gerardo Hospital, Monza, Italy
Department of Medical Sciences, Università degli Studi di Torino, Turin, Italy
Respiratory Service, Galdakao Hospital, Osakidetza Basque Health Service, Galdakao, Spain
Biocruces Bizkaia Health Research Institute, Barakaldo, Spain
Department of Infectious Diseases, E.O. Ospedali Galliera, Genova, Italy
Accident & Emergency and Emergency Medicine Unit, San Gerardo Hospital, Monza, Italy
Histocompatibilidad y Biologia Molecular, Centro de Transfusion de Madrid, Madrid, Spain
Dino Ferrari Center, Department of Pathophysiology and Transplantation, University of Milan, Milan, Italy
Neurology Unit, IRCCS Fondazione Ca’ Granda Ospedale Maggiore Policlinico, Milan, Italy
Department of Pathophysiology and Transplantation, University of Milan, Milan, Italy
Angelo Bianchi Bonomi Hemophilia and Thrombosis Center, Fondazione IRCCS Ca’ Granda Ospedale Maggiore Policlinico, Milan, Italy
Respiratory Unit, Fondazione IRCCS Ca’ Granda Ospedale Maggiore Policlinico, Milan, Italy
Department of Pathophysiology and Transplantation, Università degli Studi di Milano, Milan, Italy
Internal Medicine Department, Virgen del Rocio University Hospital, Seville, Spain
Biochemistry Department, University Hospital Vall d’Hebron, Barcelona, Spain
Centre for Medical Sciences-CISMed, University of Trento, Trento, Italy
Anesthesia and Intensive Care, Santa Chiara Hospital, APSS Trento, Trento, Italy
Department of Anesthesiology, Intensive Care and Emergency, Fondazione IRCCS Ca’ Granda Ospedale Maggiore Policlinico, Milan, Italy
Fondazione Grigioni per il Morbo di Parkinson and Parkinson Institute, ASST Gaetano Pini-CTO, Milan, Italy
School of Medicine and Surgery, University of Milano-Bico

Broad Institute of MIT and Harvard
University of California San Francisco
Institute for Molecular Medicine Finland (FIMM)
Analytic and Translational Genetics Unit
Department of Psychiatry
Veteran Affairs Connecticut Healthcare System
Yale University
Centre for Bioinformatics and Data Analysis
Department of Computational Medicine and Bioinformatics
Faculty of Industrial Sciences and Technology
Icahn School of Medicine at Mount Sinai
Genolier Innovation Network and Hub
Immediate
Clinical Research Unit of Nanoro
Sydney Brenner Institute for Molecular Biosciences
McGill University
University of Peurto Rico
Institute for Biomedical Technologies
University of Miami
Department of Biochemistry
Department of Biosciences
Department of Clinical Pathology
Department of Clinical Pathology
Neuron23
Vanderbilt University
Stroke Pharmacogenomics and Genetics
Department of Medicine
Program in Medical and Population Genetics
Department of Genetics
Departamento de Tecnología Médica
King’s College London
Naina Tech
Vrije Universiteit Amsterdam
Medical School
MNM Diagnostics
Genomics England
Heart Institute (InCor)
Department of Physiology
Montreal Heart Institute
Nantes Université
Department of Computer Science and Technology
William Harvey Research Institute
University of Helsinki
Department of Internal Medicine
Fonds de la Recherche Scientifique (FNRS) & Centre de Génétique Humaine
Department of Genetics
All India Institute of Medical Sciences Kalyani
National Laboratory of Genomics for Biodiversity
Queensland University of Technology
Department of Human Genetics
Lady Davis Institute
Kyoto-McGill International Collaborative School in Genomic Medicine
Japan Society for the Promotion of Science
Division of Life Sciences
Root Deep
Instituto do Coração
Stanford University
Department of Epidemiology
Institute for Community Medicine
GI and Liver Section
CIBEREHD
Department of Health
Rutgers Cancer Institute of New Jersey
Erasmus Medical Center
Institute of Psychiatric Phenomics & Genomics
Department of Psychiatry
Institute of Virology
Institute of Human Genetics
Department of Psychiatry
Digital Health Center
Institute of Biomedicine and Cancer Research Laboratories
IRCCS Istituto Giannina Gaslini
Department of Neurosciences
Department of Biomedical Data Science
Fundación Pública Andaluza para la Gestión de la Investigación en Salud de Sevilla
Department of Pediatrics
University Medical Center Utrecht
Instituto de Investigación Interdisciplinaria & Escuela de Medicina
Programa de Genética Humana
Department of Statistical Genetics
Department of Internal Medicine
Centro para el Desarrollo de la Investigación Científica (CEDIC)
23andMe
Washington University School Of Medicine
Department of Psychological & Brain Sciences
Institute of Biological Psychiatry
iPSYCH
Laureate Institute for Brain Research
Department of Radiology
Nork Infection Clinical Hospital
Institute of Molecular Biology NAS RA
National Research Center for Cardiac Surgery
City Infectious Disease Center
National Center for Biotechnology
Health Department of Astana
MRC Integrative Epidemiology Unit
Division of Psychitary
Institute for Molecular Medicine Finland (FIMM)
University of Liege
CHC Mont-Légia
BHUL (Liège Biobank)
Institute for Molecular Medicine Finland
Analytic & Translational Genetics Unit
Stanley Center for Psychiatric Research
CHU of Liege
University of Liege
Centre de Génétique Humaine
Service de Médecine Interne
McGill Genome Centre and Department of Human Genetics
Department of Human Genetics
Department of Twin Research
The Meakins-Christie Laboratories at the Research Institute of the McGill University Heath Centre Research Institute and Department of Medicine
Research Centre of the Centre Hospitalier de l’Université de Montréal (CRCHUM)
Centre Hospitalier de l’Université de Montréal (CHUM)
Department of Epidemiology
Department of Emergency Medicine
Emergency Department
McGill AIDS Centre
Division of Infectious Diseases
Research Centre of the Centre Hospitalier de l’Université de Montréal and Université de Montréal
School of Rehabilitation
Department of Medicine
Center for Immunity
Génome Québec
Département de Microbiologie et Infectiologie
Département de Médecine
Center for Clinical Epidemiology
Department of Neurology and Neurosurgery
Centre Intégré Universitaire de Santé et de Services Sociaux (CIUSSS) du Saguenay–Lac-Saint-Jean
Department of Pharmacology-Physiology
Division of Respiratory Medicine
Centre of Genomics and Policy
Division of Genetic Medicine
Vanderbilt Genetics Institute
Institute of Human Genetics
Institute of Genomic Statistics and Bioinformatics
Department of Gastroenterology
Department of Gastroenterology
Clinic for Cardiology
Department of Pneumology and Intensive Care Medicine
Department of Pneumology
Department of Gastroenterology
Hannover Unified Biobank
Department I of Internal Medicine
Center for Molecular Medicine Cologne (CMMC)
German Center for Infection Research (DZIF)
Institute for Human Genetics and Genomic Medicine
Department of Anesthesiology and Intensive Care Medicine
Department of Child and Adolescent Psychiatry
Department of Infectious Diseases
Department of Pneumology
Center of Human and Molecular Biology
Department of Genetics & Epigenetics
Institute of Innate Immunity
University of Toronto
Dalla Lana School of Public Health
Lunenfeld-Tanenbaum Research Institute
Simon Fraser University
University of Waterloo
The Hospital for Sick Children
Queen’s University
Mount Sinai Hospital
St Michael’s Hospital
Princess Margaret Cancer Center
Ontario Institute for Cancer Research
Dynacare Laboratory
William Osler Health System
University Health Network
Mackenzie Health
Women’s College Hospital
University of British Columbia
BC Children’s Hospital
St Paul’s Hospital
University of Calgary
Alberta Children’s Hospital
British Columbia Cancer Research Centre
Toronto General Hospital
Division of Infectious Diseases
Department of Biochemistry
The Ottawa Hospital Research Institute
University of Montreal
Ste-Justine University Health Centre
University of Toronto Ontario
MitogenDx/Eve Technologies
Genome Sciences Centre
Montreal Children’s Hospital
London Health Sciences Centre
Kidney and Metabolism Program
Sinai Health
Baycrest Health Sciences
University of Colorado
Institute of Psychiatric Phenomics and Genomics (IPPG)
Department of Genetic Epidemiology in Psychiatry
Department of Psychiatry and Behavioral Sciences
Department of Psychiatry and Behavioral Sciences
Department of Psychiatry and Psychotherapy
Institute of Virology
Department of Psychiatry and Psychotherapy
CIBER de Salud Mental
Munich Clinic Schwabing
Department of Internal Medicine II
Institute of Clinical Chemistry and Pathobiochemistry
TranslaTUM
Department of Psychiatry and Psychotherapy
German Center for Neurodegenerative Diseases (DZNE)
Neurosciences and Signaling Group
Munich Site of the German Center for Infection Research (DZIF)
Institute of Computational Biology
Department of Hygiene and Epidemiology
1st Department of Internal Medicine and Infectious Diseases Unit
Department of Epidemiology and Biostatistics
Center for Evidence-Based Medicine
University of Ioannina School of Medicine
Centre for Systems Biology
Institute of Clinical Molecular Biology
Novo Nordisk Foundation Center for Protein Research
Institut de Biotecnologia i de Biomedicina
ICREA
Institute of Biotechnology
Research Group for Evolutionary Immunogenomics
Research Unit for Evolutionary Immunogenomics
Department of Gastroenterology
Centro de Investigación Biomédica en Red en Enfermedades Hepáticas y Digestivas (CIBEREHD)
Vall d’Hebron Institut de Recerca (VHIR)
Fondazione IRCCS Ca’ Granda Ospedale Maggiore Policlinico
Department of Medicine
Microbiology Unit
Ibs.Granada Instituto de Investigación Biosanitaria
Klinik für Innere Medizin I
Emergency Department
Department for Infectious Diseases and Infection Control
Department of Medicine I
Christian Doppler Laboratory of Iron and Phosphate Biology at the Department of Medicine I
Institute of Clinical Medicine
Department of Microbiology
Hospital Clinic
European Foundation for the Study of Chronic Liver Failure (EF-CLIF)
GCAT—Genomes for Life
Genomes for Life—GCAT Laboratory
University of Milan
Department of Liver and Gastrointestinal Diseases
Digestive Diseases Unit
University of Sevilla
Instituto de Biomedicina de Sevilla (IBIS)
Hospital Universitario Virgen del Rocío de Seville
Universitat Autònoma de Barcelona
Liver Unit
Gastrointestinal Genetics Lab
Ikerbasque
Department of Liver and Gastrointestinal Diseases
Department I of Internal Medicine
University of Cologne
European Reference Network on Hepatological Diseases (ERN RARE-LIVER)
Division of Gastroenterology
Genomes for Life—GCAT lab
IRCCS Humanitas Research Hospital
Department of Biomedical Sciences
Zentrum für Humangenetik Regensburg
University Hospital Schleswig-Holstein (UKSH)
Section for Gastroenterology
Research Institute for Internal Medicine
Norwegian PSC Research Center
Private University in the Principality of Liechtenstein
Division of Rheumatology
Division of Genetics
Department of Biomedical Informatics
Center for Data Sciences
Institut de Biotecnologia i de Biomedicina
Institute for Medical Microbiology and Hospital Epidemiology
Randaberg Municipality
Department of Quality and Health Technology
University Hospital Schleswig-Holstein
Research Department
Department of Genetics and Bioinformatics (HDGB)
Centre for Genetics and Genomics Versus Arthritis
Department of Intensive Care
Clinical Biochemistry Department
Research Center and Memory Clinic
CIBERNED
Department of Biochemistry
Department of Acute Medicine
Division of Neurogenetics and Molecular Psychiatry
Department of Neurodegenerative Diseases and Geriatric Psychiatry
German Center for Neurodegenerative Diseases (DZNE)
Department of Psychiatry
Department of Anesthesiology and Intensive Care
Pediatrics and Tettamanti Center
Biochemistry Department
Centre for Multidisciplinary Research in Health Science (MACH)
Gastroenterology Unit
Microbiology Unit
Department of Infectious Diseases
Humanitas Gavazzeni-Castelli
Microbiology Department
Department of Respiratory Diseases
Department of Respiratory Medicine and Allergology
Centro de Investigación Biomédica en Red de Enfermedades Respiratorias (CIBERES)
Centro de Investigación Biomédica en Red de Epidemiología y Salud Pública (CIBERESP)
Consejo Superior de Investigaciones Científicas
Department of Infectious Diseases
Department of Transfusion Medicine and Haematology Laboratory
Respiratory Medicine & International Health
Division of Clinical Infectious Diseases
Clinical Tuberculosis Unit
Internal Medicine Department
Humanitas Clinical and Research Center
Respiratory Service
Department of Clinical and Molecular Medicine
Department of Medicine
Department of Internal Medicine II
Department of Anesthesiology and Critical Care
Immunohematology Department
European Reference Network on Hepatological Diseases (ERN RARE-LIVER)
Department of Medical Sciences
Respiratory Service
Biocruces Bizkaia Health Research Institute
Department of Infectious Diseases
Accident & Emergency and Emergency Medicine Unit
Histocompatibilidad y Biologia Molecular
Dino Ferrari Center
Neurology Unit
Department of Pathophysiology and Transplantation
Angelo Bianchi Bonomi Hemophilia and Thrombosis Center
Respiratory Unit
Department of Pathophysiology and Transplantation
Internal Medicine Department
Biochemistry Department
Centre for Medical Sciences-CISMed
Anesthesia and Intensive Care
Department of Anesthesiology
Fondazione Grigioni per il Morbo di Parkinson and Parkinson Institute
School of Medicine and Surgery

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